BIO 401. Life at the Edge: The Physiological Ecology of Extreme Environments.

      Dr. Bruce W. Grant and Dr. Itzick Vatnick
      Department of Biology, Widener University, Chester, PA, 19013
      BWG office Loveland Hall # 9, ext. 4017,
      IV office Kirkbride # 516, ext. 4245
      grant@pop1.science.widener.edu and
      vatnick@pop1.science.widener.edu
      classes meet: MWF 1:00-1:50, Kirkbride 500.
      project lab: Tuesday 11:00-4:45, Kirkbride 503 and Kirkbride 420.
      4 credit hours.

      • Current Assignments.
      • Course Objectives and Description.
      • Class Schedule.
      • Physiological Ecology Research and Educational Resources.
      • Please Read This Comment.



      Efferia texana, Big Bend National Park, Texas
      photo © BW Grant, 1997
      Page last updated: 11 November 2003 and has been accessed times since 1 September 1997. We thank Widener University (particularly Larry Panek, Marc Brodkin, and Marty Schultz) for server access and technical support. We also thank Dr. David Coughlin who co-taught this course with us in previous years, and who continues to provide essential consulting and mentoring services for some of the students' course research projects.



      Course Objectives and Description.

      (1) You will better understand the evolutionary adaptation of physiology to environment, i.e., the problems that the biophysical extremes of this world impose upon life and the solutions that have evolved in response.

              This course will deal with the evolutionary adaptation of physiology to environment (e.g., temperature, water, light, salinity, pressure, nutrients, and toxins). Students will learn how individuals interact with their biophysical environment, and how physiological processes offer integrated responses to environmental stress. Examples will illustrate adaptation to warm desert environments, arctic and alpine environments, freshwater and saline wetlands, deep sea environments, and human-disturbed environments. Although topics represent the extremes of this world, an important message is that over evolutionary time all environments are variable and the evolutionary rules affecting physiology at the extremes apply everywhere life is always at the edge.

              Underlying themes of this course are the similarity of solutions to common problems (such as adaptations to water stress in deserts and saline wetlands) and the multiplicity of solutions evolved among life forms to the same environmental stress (such as the diversity of temperature regulation modes in thermally heterogeneous environments). We will primarily focus the course on animals; however, moneran, protistan, fungal, and plant adaptations to environmental stress are also excellent sources of independent research projects (described below) and we encourage students to pursue whatever interests them.

              A major venue for the teaching and learning of these basic principles of physiological design will be a seminar-style weekly discussion class of primary literature from the field of physiological ecology. For this, you must locate, interpret, and critically analyze published journal articles relevant to the course topics. More details on this will be presented in class.


      (2) You will understand the process of physiological inquiry which is the scientific method, through which natural phenomena are observed, interpreted, and reported. Through the scientific method one gains an understanding of our natural world and the effects of human activities upon it.

              A major portion of this course is dedicated to student-designed independent research projects involving laboratory experiments on any topic of the your interest in physiological ecology, and using available equipment funded by an NSF ILI grant to Dr's. Vatnick and Grant. You will select a research topic, make an oral proposal presentation, conduct the research, and present an oral talk and poster at in-class research symposia toward the end of the semester. In addition, you must locate and interpret background literature relevant to each your research project and lead a group discussion of your literature during one of the weekly lab meetings/ seminars.


      (3) You will improve your cognitive skills at critical thinking and reflective judgement. Through the processes of designing, conducting, and presenting your own course research project, and of reading and critically analyzing published research results to understand the basic ecological and evolutionary principles of physiological design, you will be using and improving your higher level thinking and information processing skills.

              These are the kinds of "good thinking" skills that are essential in grappling with "problems for which no clear-cut solution can be identified by using only inductive or deductive logic; solutions to these problems involve other characteristics of reasoning." (King and Kitchener 1994). In this course, you will construct your own understanding of physiological phenomena (as constrained, facilitated, and calibrated by us). Through this process, you will be challenged by complex, multifaceted, novel, and unexpected conceptual, scientific, educational, social, and technical problems. This is what research is all about. You'll need to use your head to solve these problems. Intellectual growth will result.






      Course Requirements.

            Attendance is required and will benefit your grade. Missed in-class assignments, quizzes, etc. cannot be made up unless extraordinary circumstances are documented for one's absense.

            The principal text is by Knut Schmidt-Nielsen. 1997. Animal Physiology: Adaptation and Environment. Cambridge University Press. 5th Edition.

            This text is THE classic text in this discipline. It emphasizes the literature and includes excellent summaries of what is known in many important areas of physiological ecology. In addition, Schmidt-Nielsen’s book creatively weaves into the narrative important applications of ecological knowledge to real world environmental problem solving. Required readings and www activities will be noted in class.

            Since this course includes an intensive writing component (essay-based exams, a major research paper, etc.), an additional text I require you to access is the Elements of Style by Strunk and White. This is one of the best books on English usage ever written, and should be one of the most important books you will buy (or bookmark) and read as a student. Note: if you want to access a web-based version of this text for free press http://www.bartleby.com/141/index.html.






      Friday Seminars.

            During this semester, we will devote the Friday 1:00 class periods to seminar-style discussions on topics of relevance to this course. We will lead the first one of these to give you the idea about the format, and then you will take turns leading the others during the rest of the semester. For each seminar, the seminar leader(s) will select a research paper, review, etc., and we will copy and distribute it to the class. You should sign up for a seminar date and topic as soon as you can.





      Course Assessment.

            There will be a midterm and a final project exam (no FINAL exam), weekly assignments, and weekly lab meetings/seminars at which research papers will be discussed. A detailed breakdown of the components of assessment in this course appears below:

      Midterm Exam150 points
      Seminars on Reading Sets150 points
      Research Project Proposal50 points
      Project Results-to-Date Powerpoint Presentations   50 points
      Weekly Assignments100 points
      Review of Peer's Poster50 points
      Final Project Poster150 points
      Project Exam150 points
      Data Management150 points
      TOTAL   1000 points





      Fall 2003, Class Schedule.

      class # Lab # Sem # datetopic
      #1Fri, Sept 5What is physiological ecology?
      #1Mon, Sept 8Discussion on perspectives of physiological ecology.
      #1Tue, Sept 9Principles of Organismal Biophysical Design: Measuring and Modeling the Theory of Cooling
      #2Wed, Sept 10Biophysical Ecology: Plant and Animal Heat Budgets (Chapters 6 & 7)
      #2Fri, Sept 12Seminar on Reading Set 1
      #3Mon, Sept 15Animal Energy and Mass Budgets (Chapter 4, pp 129-163)
      #2Tue, Sept 16Principles of Organismal Biophysical Design: Modeling and Measuring Lizard Operative Environmental Temperatures
      #4Wed, Sept 17Animal Metabolism
      #3Fri, Sept 19Seminar on Reading Set 2
      #5Mon, Sept 22Animal Metabolism (Chapter 5, pp. 170-179, 192-200)
      #3Tue, Sept 23Animal Metabolism - Lab 1
      #6Wed, Sept 24Animal Metabolism - Oxygen Supply (Chapter 1)
      #4Fri, Sept 26Seminar on Reading Set 3
      #7Mon, Sept 29Animal Metabolism - Blood (Chapter 2)
      #4Tue, Sept 30Animal Metabolism - Lab 2
      #8Wed, Oct 1Animal Metabolism - Circulation (Chapter 3)
      #5Fri, Oct 3Seminar on Reading Set 4
      #9Mon, Oct 6Project Proposal Presentations
      #5Tue, Oct 7project work day
      #10Wed, Oct 8Animal Metabolism - Excretion (Chapter 9)
      #6Fri, Oct 10Seminar on Reading Set 5
      #11Mon, Oct 13project work day
      #6Tue, Oct 14Midterm Exam
      #12Wed, Oct 15project work day
      Fri, Oct 17*** fall break ***
      Mon, Oct 20*** fall break ***
      #7Tue, Oct 21project work day
      #13Wed, Oct 22project work day
      #7Fri, Oct 24Seminar on Reading Set 6
      #14Mon, Oct 27project work day
      #8Tue, Oct 28project work day
      #15Wed, Oct 29project work day
      #8Fri, Oct 30Seminar on Reading Set 7
      #16Mon, Nov 3project work day
      #9Tue, Nov 4project work day
      #17Wed, Nov 5project work day
      #9Fri, Nov 7Seminar on Reading Set 8
      #18Mon, Nov 10project work day
      #10Tue, Nov 11project work day
      #19Wed, Nov 12project work day
      #10Fri, Nov 14Seminar on Reading Set 9
      #20Mon, Nov 17project work day
      #11Tue, Nov 18Project Results-to-Date Powerpoint Presentations
      #21Wed, Nov 19project work day
      #11Fri, Nov 21Seminar on Reading Set 10
      #22Mon, Nov 24project work day
      #12Tue, Nov 25project work day
      #23Wed, Nov 26project work day
      Fri, Nov 28*** thanksgiving ***
      #24Mon, Dec 1project work day
      #13Tue, Dec 2project work day
      #25Wed, Dec 3project work day
      #12Fri, Dec 5Seminar on Reading Set 11
      #26Mon, Dec 8project work day
      #14Tue, Dec 9project work day
      #27Wed, Dec 10project work day
      #13Fri, Dec 12Final Project Posters and Project Exams Due





      Physiological Ecology Projects Students Have Done.
        ... under construction.





      Physiological Ecology Research and Educational Resources.
        ... under development...





      Please Read This Comment.
              You are welcome to download some or all of the material we have posted at this site for your use in your physiological ecology course. This does not include commercial uses for profit. If you do use any lengthy exerpts (more than 2 lines) of the material above, we request that you formally acknowledge this site and/or sites we have acknowledged as the source(s). We also request that you reciprocate and send us a copy of your physiological ecology educational materials so that we may see what you have put together.

        Please send comments or suggestions to Bruce W. Grant: grant@pop1.science.widener.edu.

        Copyright - Bruce W. Grant and Itzick Vatick, 2003.

        "The views expressed here are those of the page authors. Widener University does not review individual pages of faculty, administrators, or students and will not take responsibility for them."

      PLEASE DO NOT PRINT OUT THIS WEB SITE !!!