Biology 313. Evolution.

      Dr. Bruce W. Grant
      Department of Biology, Science Division, Widener University
      Chester, PA, 19013, BWG office K414/K418 x4017
      grant@pop1.science.widener.edu
      classes meet: MWF 1:00-1:50, Tue 2:00-5:00, Kirk. 313.
      4 credit hours.

      • Current Assignments.
      • Course Objectives.
      • Course Structure.
      • Class Schedule.
      • Handouts Available on the Web.
      • Projects Students Have Done.
      • Evolution Research and Educational Resources.
      • Please Read This Comment.

      • To download a *.PDF (167Kb) of this file press → here.



      Page last updated: 15 January 2007 and has been accessed times since 1 September 1997. I thank Widener University (particularly Harry Augensen and Marty Schultz) for server access and technical support. I also thank Dr.'s Itzick Vatnick and David Coughlin with whom I co-designed and co-taught this course in previous years.



      Course Objectives.

             Evolution is the most powerful and elegant theory in biology. Without evolution, biology becomes a disparate set of technical sub-disciplines. Evolutionary explanations pervade all fields in biology and bring them together under one theoretical umbrella. For this reason, evolution is the unifying concept of the biological sciences.

      There are three major objectives of this course:

      • (1) Increase Your Understanding of the Causes, Processes, and Consequences of Evolution. This is the domain related to disciplinary content – the “knowing things” domain. This objective includes three sub categories:

          (a) Understand the Principal Mechanisms of Evolution. You will understand how the mechanisms of microevolutionary change (genetic drift, gene flow, mutation, and natural selection) create, maintain, and destroy genetic variation and have resulted in the remarkable diversity of life forms present today. Also, you will learn that these microevolutionary mechanisms affect ongoing ecological processes and can explain many current interactions among organisms.

          (b) Understand the Major Evolutionary Trends in Biotic Diversity. Several major evolutionary trends occurred during the history of life on Earth (e.g. often increasing complexity – but not always, evolution of predation, colonization of land, etc.). You will understand (i) what constitutes a major evolutionary trend, (ii) the concept of macroevolution, (iii) the role of natural selection in molding key traits, and (iv) the pathways along which life forms “progressed” within the major phyletic lineages.

          (c) Understand the Evolutionary Origin of Life. A common evolutionary ancestry interconnects all life forms on Earth. You will understand the theory of how life evolved on Earth and diversified into the variety of forms we see today.

      • (2) Improve Your Scientific Literacy About How Evolutionary Knowledge is Constructed Using the Scientific Method. It is through the process of scientific inquiry using the scientific method that natural phenomena are observed, interpreted, and reported. Science is a “way of knowing” about the world around us with which one gains an understanding of our environment and the effects of human activities upon it. This is the domain related to knowing how to construct the disciplinary content knowledge from (1) – thus, this objective spans both the “knowing how to do things” and “knowing how to know things” domains. This objective includes three sub categories:

          (a) Understand How to Investigate and Model the Dynamics of Evolutionary Change in Ecological Systems (Populations and Communities). You will improve your skills at applying the scientific method to investigating, modeling, and understanding evolution in simulated populations and communities.

          (b) Understand the Principles of Phylogenetic Systematics and the Methods of Bioinformatics. Bioinformatics is one of the fastest growing fields of biology. Huge advances are being made as entire genes and genomes are being sequenced and shared. Although the principal goal of many of these efforts is in applied biomedical, agricultural, or environmental research, exciting and new insights are emerging about evolutionary relationships among organisms at all taxonomic levels. Previous phylogenies, based mostly on morphological or developmental data and ill-defined guesswork, are crumbling almost daily beneath this onslaught of hi-resolution rigorous molecular genetic analyses. In this course, you will understand these techniques, and draw from the vast internet database of published genetic research and pose and text hypotheses about evolutionary relationships among organisms of interest to you.

          (c) To Improve Your Scientific Thinking Skills Including Systems Thinking and Reflective Judgment. Systems thinking approaches complex problems from an holistic rather than a reductionist perspective. Reflective judgment extends evidence-based thinking to the analysis of problems such as the complex challenges facing the sustainability of our global society today. These skills will be developed throughout all activities in this course.

      • (3) Understand the Uses and Effects of the Theory of Evolution in Society. Evolutionary knowledge is at the fore-front of the biological revolution that has characterized the past several decades, and will likely dominate biology, medicine, and policy for the rest of this century. For example, biotechnology is now being used to manipulate the genome to improve our domesticated plants and animals (genetic engineering), our environment (bioremediation), and ourselves (numerous desirable examples such as gene therapy, but some much less so such as eugenics). In addition, evolutionary theories have been and continue to be brought into social debates to sometimes explain social behavior (sociobiology) but often to justify racism, social injustice, or even cultural genocide. Finally, recently the theory of evolution has come under vigorous attack by extremists who proclaim themselves to be “creation scientists” or proponents of "intelligent design" and demand access to the minds of our youth through public school science curricula. No course in evolution would be complete without an exploration of the non-scientific belief system that these critics advocate.



      Course Structure.

        Requirements. Attendance is required and will benefit your grade. There will be weekly journal clubs (Fridays), several presentations, two in-class exams and a final exam based on these two exams, an individualized project-based "midterm exam" (dates on the next page, based on your choice of course project), and numerous other in-class writing assignments and quizzes. All of the journal club assignments and your course project will involve cycles of revision; consequently, this course is designated "Writing Enriched."

               There is NO one required text. I will draw from several published texts, copies available in the Evolutionary Ecology Reading Room (K414); however, a great deal will come from Freeman and Herron (2007). Evolutionary Analysis. 4th edition. Prentice Hall. This is an excellent text by two of the leaders in the field of evolutionary biology. This text blends an unique combination of natural history, rigorous theory, and critical experimentation using current literature. It excellently summarizes what is known in many important areas of evolutionary biology using crisp and simple prose and great examples. Required readings will be announced in class.

               In addition, for the unit on phylogenetic systematics, there is a workbook: Hall, B. G. 2004. Phylogenetic Trees Made Easy: A How-To Manual for Molecular Biologists. 2nd edition, Sinauer. Copies will also be available in K414.

               Lastly, I will be assigning chapters in Jonathan Weiner’s Pulitzer Prize winning book – “The Beak of the Finch” to improve your understanding of Darwinian natural selection and the effort required in field work to detect it.
           


        Assessment. Your grades will be based upon weekly journal club writing assignments using the primary literature in evolution, several projects with presentations and writing assignments, in-class exams, an individualized "Project Exam," and a Final Exam during final exam week. Details are posted on Campus Cruiser. Lastly, to encourage and reward participation, there will be frequent in-class assessment and evaluation activities. Your final grade is based on your total accumulated points, and if everyone earns >90%, then everyone gets an “A.”





      Syllabus (Spring 2007)

        class #datetopic
         
         Evidence for Evolution
        labTue Lab, Jan 16Introduction to Evolution
        #1Wed, Jan 17Misconceptions About Evolution
        #2Fri, Jan 19Journal Club 1 - Why Teach Evolution?
         Microevolution
        #3Mon, Jan 22Introduction to Microevolution
        labTue Lab, Jan 23Modeling Mutation
        #4Wed, Jan 24Modeling Selection
        #5Fri, Jan 26Journal Club 2 - Detecting Natural Selection
        #6Mon, Jan 29Modeling Genetic Drift 1
        labTue Lab, Jan 30Microevolution Mini-projects Begin
        #7Wed, Jan 31Microevolution Mini-project Work Day
        #8Fri, Feb 2Journal Club 3 - Microevolution
        #9Mon, Feb 5Microevolution Mini-project Work Day
        labTue Lab, Feb 6Microevolution Mini-project Presentation Day
        #10Wed, Feb 7Quantitative Genetics and Phenotypic Plasticity
        #11Fri, Feb 9Journal Club 4 - Conservation Genetics - 1
        #12Mon, Feb 12EXAM 1
        labTue Lab, Feb 13Evolution in Kin, Groups, Lineages, and Clades
        #13Wed, Feb 14Sexual Selection
        #14Fri, Feb 16Journal Club 5 - Altruism and Group Selection
         On to Macroevolution
        #15Mon, Feb 19Coevolution Among Competitors
        labTue Lab, Feb 20Modeling Coevolution - Hosts/Parasites
        #16Wed, Feb 21Coevolution Among Predators and Prey
        #17Fri, Feb 23Journal Club 6 - Coevolutionary Ecology
        #18Mon, Feb 26Coevolution Among Mutualists
        labTue Lab, Feb 27Workshop on the Evolution of Humans
        #19Wed, Feb 28Origin of Species
        #20Fri, Mar 2Journal Club 7 - Mass Extinction
         Mon, Mar 5spring break
         Tue Lab, Mar 6spring break
         Wed, Mar 7spring break
         Fri, Mar 9spring break
        #21Mon, Mar 12Origin of the Universe and Our Solar System
        labTue Lab, Mar 13Evidence for Terrestrial Origins to Life on Earth
        #22Wed, Mar 14Evidence for the Origins of Biodiversity
        #23Fri, Mar 16Journal Club 8 - Venus, Mars, Titan, and Europa
         Bioinformatics and Phylogenetic Inference
        #24Mon, Mar 19Phylogenetic Inference
        labTue Lab, Mar 20Phylogenetic Inference Mini-projects Begin
        #25Wed, Mar 21Phylogenetic Inference Mini-project Work Day
        #26Fri, Mar 23Journal Club 9 - Molecular Clocks
        #27Mon, Mar 26Phylogenetic Inference Mini-project Work Day
        labTue Lab, Mar 27Phylogenetic Inference Presentation Day
        #28Wed, Mar 28Course Project Selection Day
        #29Fri, Mar 30Journal Club 9 - TBA
        #30Mon, Apr 2EXAM 2
        labTue Lab, Apr 3Course Project Work Day - Proposals Due
        #31Wed, Apr 4Topics in Evolutionary Ecology - TBA
        #32Fri, Apr 6Spring Holiday
        #33Mon, Apr 9Topics in Evolutionary Ecology - TBA
        labTue Lab, Apr 10Course Project Work Day
        #34Wed, Apr 11Topics in Evolutionary Ecology - TBA
        #35Fri, Apr 13Journal Club 10 - TBA
        #36Mon, Apr 16Topics in Evolutionary Ecology - TBA
        labTue Lab, Apr 17Course Project Work Day
        #37Wed, Apr 18Topics in Evolutionary Ecology - TBA
        #38Fri, Apr 20University-Wide Undergraduate Projects Day
        #39Mon, Apr 23Course Project Work Day - "PROJECT EXAMS" begin
        labTue Lab, Apr 24Course Project Work Day
        #40Wed, Apr 25Course Project Work Day
        #41Fri, Apr 27Journal Club 11 - TBA
        #42Mon, Apr 30Course Project Work Day - "PROJECT EXAMS" due
        labTue Lab, May 1Final Project Presentations
        #43Wed, May 2Revisions of "PROJECT EXAMS" Due

        FINAL EXAM date and time TBA during Final Exam Week.

        Class schedules and exams from previous years:

        • Spring 2005, Class Schedule and the Final Exam
        • Fall 2003, Class Schedule
        • Fall 2001, Class Schedule
        • Fall 1999, Class Schedule, Exam 1, Exam 2, Exam 3, and the Final Exam
        • Fall 1997, Class Schedule


      Handouts Available on the Web.

      *** under construction ***
      Projects Students Have Done.

      *** under construction ***


      Evolution Research and Educational Resources.

      *** under construction ***


      Please Read This Comment.
              You are welcome to download some or all of the material we have posted at this site for your use in your evolution course. This does not include commercial uses for profit. If you do use any lengthy exerpts (more than 2 lines) of the material above, we request that you formally acknowledge this site and/or sites we have acknowledged as the source(s). We also request that you reciprocate and send us a copy of your evolution educational materials so that we may see what you have put together.

        This page was written by Bruce W. Grant, grant@pop1.science.widener.edu.

        Copyright - Bruce W. Grant, 2007.

        "The views expressed here are those of the page authors. Widener University does not review individual pages of faculty, administrators, or students and will not take responsibility for them."

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